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Sicyos pachycarpus

Hook. & Arn. (1832)

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-Rosids Order: Cucurbitales Family: Cucurbitaceae Genus: Sicyos

‘ānunu [anunu], kūpala [kupala]

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Description

Key Characters:

Growth Form: Climbing vines, perhaps with perennial rootstock.

Stems: Stems slender, up to several m long, ca. 1 mm in diameter, glabrous.

Roots:

Leaves: Leaves simple. Alternate. Blades broadly ovate–cordate or reniform with a broad basal sinus, 7–26 cm long, 7–28 cm wide, usually shallowly 5–7–lobed or angled, rarely deeply 5–7–lobed, the lobes usually broadly triangular, rarely narrowly obovate and lobulate. Apex usually obtuse and mucronate, the central lobe abruptly acuminate, the slender apex often breaking early. Surfaces glabrous or Upper surfaces sparsely scabrid; lower surfaces scabrid with short white hairs. Margins denticulate. Petioles 1.5–3 cm long. Stipules absent; tendrils in the stipular position 2–branched.

Flowers: Flowers in staminate flowers in pubescent or puberulent panicles 3–12 cm long, peduncles 2–9 cm long, pedicels 2–3 mm long; pistillate flowers (5–)10–50 per head, sessile on peduncles 0.4–2(–4) cm long. Flowers usually unisexual (and the plants monoecious or dioecious), actinomorphic. Calyx of 5 sepals; sepals inserted on hypanthium. Corolla of staminate flowers greenish yellow, 5-lobed, 3–5 mm in diameter, petals inserted on hypanthium alternate with calyx lobes, usually connate at base, the limb often rotate; Corolla of pistillate flowers 3–5-lobed, perianth usually smaller than in staminate flowers. Staminate flowers: anthers coherent, rarely spreading. Staminate flowers: Stamens (2)3(–5), inserted at base of hypanthium; filaments connate into a column; anthers curved or flexuous, coherent, rarely spreading, opening by longitudinal slits; pistillate flowers: staminodes absent. Ovary inferior, of a single carpel or 3-carpellate with carpels connate into a single 3-lobed cell; ovule 1, placentation parietal; stigma shortly 3-lobed, the lobes recurved.

Fruit: Fruit brown or black; ovoid; attenuate; 3–8 mm long (incl. beak); compressed or angled toward base; glabrous or puberulent; abruptly contracted into a short beak; indehiscent. Seeds 1 per fruit; ovate; endosperm absent.

Ploidy: 2n = 24*

Habitat: Widespread in herb or shrubland coastal communities; dry forest; and alien vegetation such as Leucaena or Prosopis shrubland; on coral sand or clay loam; primarily on lower leeward slopes.

Elevation Range: 0–900 m.

Historical Distribution

Images

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Synonyms (32)

Sicyocarya atrimaculata H.St.John (1987)
Sicyocarya rostrata H.St.John (1987)
Sicyos laysanensis H.St.John (1970)
Sicyos microcarpus H.Mann (1867)
Sicyos microcarpus subsp. kanepuuensis H.St.John (1987)
Sicyos microcarpus subsp. lanaiensis H.St.John (1987)
Sicyos microcarpus subsp. laysanensis (H.St.John) H.St.John (1987)
Sicyos microcarpus subsp. microcarpus
Sicyos microcarpus subsp. puberulus H.St.John (1987)
Sicyos microcarpus var. ewaensis H.St.John (1987)
Sicyos microcarpus var. forbesii H.St.John (1987)
Sicyos microcarpus var. haleakalaensis H.St.John (1987)
Sicyos microcarpus var. kalaupapaensis H.St.John (1987)
Sicyos microcarpus var. kamoaensis H.St.John (1987)
Sicyos microcarpus var. kauensis H.St.John (1987)
Sicyos microcarpus var. kealiaensis H.St.John (1987)
Sicyos microcarpus var. kilaueaensis H.St.John (1987)
Sicyos microcarpus var. koeleensis H.St.John (1987)
Sicyos microcarpus var. koloaensis H.St.John (1987)
Sicyos microcarpus var. lanaiensis H.St.John (1987)
Sicyos microcarpus var. laysanensis (H.St.John) H.St.John (1987)
Sicyos microcarpus var. meeboldii H.St.John (1987)
Sicyos microcarpus var. obovoideis H.St.John (1987)
Sicyos microcarpus var. oniniensis H.St.John (1987)
Sicyos microcarpus var. paneensis H.St.John (1987)
Sicyos microcarpus var. pohakeaensis H.St.John (1987)
Sicyos microcarpus var. trilobata H.St.John (1987)
Sicyos microcarpus var. woolfordii H.St.John (1987)
Sicyos niger H.St.John (1987)
Sicyos nihoaensis H.St.John (1970)
Sicyos pachycarpa Hook. & Arn.
Sicyos remyanus Cogn. (1881)

Uses and Culture

USES

Natural History

Statewide Status

Endemic

Island Status

Ni'ihau(Incl. Lehua) Endemic
Kaua'i Endemic
O'ahu Endemic
Molokai Endemic
Kamole(Laysan Island) Endemic
Lana'i Endemic
Kaho'olawe Endemic
Maui Endemic
Hawai'i Extirpated
Nihoa Endemic

Dispersal Agents


Pollinators

Bibliography

Name Published In: Bot. Beechey Voy.: 83 (1832)

Other References

Wagner et al. 1990:579 (La, N, Ni, K, O, Mo, L, M, Ka, H); Faccenda, Arthur, et al. 2025:168 (remove Ni [no evidence of presence]); Munro 418 BISH (L†, 1929); Matt Keir pers. comm. 2025/Lorence 7991 BISH (H†, 1997)

Occurrences

SNo. Scientific Name Locality Habitat Basis of Record Description Recorded By Record Number Island Source Date
1 Sicyos pachycarpus 1st small gulch NW of Puu Pane Preserved_Specimen Degener, O. 21363 Oahu BISH 1950-04-02
2 Sicyos pachycarpus East Maui, Hana District, Nuu Preserved_Specimen Vine climbing over 8 m Prosopis trees; lvs membranous, dark green above, pale green below. Male fl white, female fl white, utricles green then black. Stem 5-angled, herbaceous, green. St.John, H. 26771 Maui BISH 1971-03-28
3 Sicyos pachycarpus Mountains near Koele Preserved_Specimen Large vine. Forbes, C.N. 95 Lanai BISH 1913-06-01
4 Sicyos pachycarpus Kanaio, S slope of Haleakala Preserved_Specimen stems- 5-sided, the ribs whitish, leaf underside like sandpaper, flowers greenish white. Forbes, C.N. 1821 Maui BISH 1920-03-03
5 Sicyos pachycarpus Laysan Preserved_Specimen Herbst, D.R. 9063 Laysan Island BISH 1988-06-25
6 Sicyos pachycarpus Waialua, Kaena, Manini Pali Preserved_Specimen vine Herbst, D.R. 5844 Oahu BISH 1976-04-25
7 Sicyos pachycarpus Puehu Rd Preserved_Specimen In flower and early fruit - Mar. 6 1979. Hobdy, R.W. 435 Molokai BISH 1979-03-15
8 Sicyos pachycarpus East Maui, Makawao District, Mauka of Puu O Kali hill Preserved_Specimen Fruits very dark when mature. Hobdy, R.W. 2063 Maui BISH 1984-04-19
9 Sicyos pachycarpus Small gulch S of the larger unnamed gulch behind the Waikapu Rock Crusher Preserved_Specimen Hobdy, R.W. 1101 Maui BISH 1981-05-22
10 Sicyos pachycarpus Nanakuli Valley below Mauna Kapu Preserved_Specimen Degener, O. 17415 Oahu BISH 1935-12-01
11 Sicyos pachycarpus East Maui, Hana District, Nuu Preserved_Specimen Vine climbing over 8 m Prosopis trees; lvs membranous, dark green above, pale green below. Male fl white, female fl white, utricles green then black. Stem 5-angled, herbaceous, green. St.John, H. 26771 Maui BISH 1971-03-28
12 Sicyos pachycarpus Nu`u along roadside in pasture Preserved_Specimen Differs from description of taxon in the stems being>1mm and not completely glabrous. Oppenheimer, H.L. 30905 Maui BISH 2009-03-13
13 Sicyos pachycarpus Preserved_Specimen lge lvs & almost mature fruits Kepler, A.K. 9223 Laysan Island BISH 1992-04-16
14 Sicyos pachycarpus Kamao Preserved_Specimen Munro, G.C. 418 Lanai BISH 1929-05-16
15 Sicyos pachycarpus East Maui, Makawao District, Auwahi, along Piilani Hwy above and E of Pimoe Preserved_Specimen Corolla white; leaves lobed to deeply parted, sometimes on same individual. Wagner, W.L. 5807 Maui BISH 1988-03-04
16 Sicyos pachycarpus Below Haleakala Ranch pipeline Preserved_Specimen Long trailing vine. Munro, G.C. 801 Maui BISH 1928-02-20
17 Sicyos pachycarpus Mountains near Koele Preserved_Specimen Large vine. Forbes, C.N. 95 Lanai BISH 1913-06-01
18 Sicyos pachycarpus N side of ridge, 1st ridge S of Kanehoa, Waianae Mts., Honouliuli Preserved_Specimen vine 3 m long Kikudome, G.Y. 431 Oahu BISH 1951-05-27
19 Sicyos pachycarpus West Maui, Makawao, Auwahi, Piilani Hwy., above and E of Pimoe Preserved_Specimen Corolla white; leaves lobed to deeply parted, sometimes on same individual. Wagner, W.L. 5807 Maui BISH 1988-03-04
20 Sicyos pachycarpus East Maui, Hana District, Nuu Preserved_Specimen Vine climbing over 8 m Prosopis trees; lvs membranous, dark green above, pale green below. Male fl white, female fl white, utricles green then black. Stem 5-angled, herbaceous, green. St.John, H. 26771 Maui BISH 1971-03-28
21 Sicyos pachycarpus Waianae Mts., Kealia Trail above Dillingham Airfield Preserved_Specimen male flowers white Wagner, W.L. 5319 Oahu BISH 1984-03-02
22 Sicyos pachycarpus W. slope of Tanager Peak Preserved_Specimen Christophersen, E. 9 Nihoa BISH 1924-07-12
23 Sicyos pachycarpus Mountains near Koele Preserved_Specimen Large vine. Forbes, C.N. 95 Lanai BISH 1913-06-01
24 Sicyos pachycarpus Kalulu Preserved_Specimen Munro, G.C. 535 Lanai BISH 1916-03-29
25 Sicyos pachycarpus Kaluakauila Valley, Keawaula, Waianae Mts on Grevillea trunks, partly sunny, dry, well sloped Preserved_Specimen Obata, J.K. 283 Oahu BISH 1975-04-06
26 Sicyos pachycarpus firebreak trail between Huliwai and Ekahanui Gulch Preserved_Specimen 5 m long Kikudome, G.Y. 432 Oahu BISH 1951-05-27
27 Sicyos pachycarpus East Maui, Hana District, Nuu Preserved_Specimen Vine climbing over 8 m Prosopis trees; lvs membranous, dark green above, pale green below. Male fl white, female fl white, utricles green then black. Stem 5-angled, herbaceous, green. St.John, H. 26771 Maui BISH 1971-03-28
28 Sicyos pachycarpus Kalulu Preserved_Specimen Munro, G.C. 535 Lanai BISH 1916-03-24
29 Sicyos pachycarpus East Maui, Makawao District, Auwahi, along Piilani Hwy above and E of Pimoe Preserved_Specimen Corolla white; leaves lobed to deeply parted, sometimes on same individual. Wagner, W.L. 5807 Maui BISH 1988-03-04
30 Sicyos pachycarpus Ridge top of N Palawai Napepeiauolelo Ridge (just N of Palikea), S Waianae Range Growing under a canopy of Eucalyptus but growing over a juvenile Eucalyptus tree on a dry ridge top. Several plants scattered in the vicinity. Preserved_Specimen Obata, J.K. 428 Oahu BISH 1980-03-30
31 Sicyos pachycarpus Kealia trail Preserved_Specimen Takeuchi, W.N. 61 Oahu BISH 1984-02-10
32 Sicyos pachycarpus Makaha Valley Preserved_Specimen trailing or climbing vine Kores, P.J. Oahu BISH 1979-05-12
33 Sicyos pachycarpus Laysan Preserved_Specimen White blossom Bryan, W.A. Laysan Island BISH 1911-05-01
34 Sicyos pachycarpus Makaha Valley Preserved_Specimen trailing or climbing vine Kores, P.J. Oahu BISH 1979-05-12
35 Sicyos pachycarpus East Maui, Hana District, Nuu Preserved_Specimen Vine climbing over 8 m Prosopis trees; lvs membranous, dark green above, pale green below. Male fl white, female fl white, utricles green then black. Stem 5-angled, herbaceous, green. St.John, H. 26771 Maui BISH 1971-03-28
36 Sicyos pachycarpus East Maui, Hana District, Nuu Preserved_Specimen Vine climbing over 8 m Prosopis trees; lvs membranous, dark green above, pale green below. Male fl white, female fl white, utricles green then black. Stem 5-angled, herbaceous, green. St.John, H. 26771 Maui BISH 1971-03-28
37 Sicyos pachycarpus West side of coral plains at Ewa, near Barber's Point, ocean side of Malakole Rd. near Camp Malakole Military Reservation Preserved_Specimen Nagata, K.M. 1656 Oahu BISH 1978-06-21
38 Sicyos pachycarpus Coral plain below Ewa and Sisal Preserved_Specimen Forbes, C.N. 2323 Oahu BISH 1916-03-14
39 Sicyos pachycarpus Kaena Preserved_Specimen Munro, G.C. 338 Oahu BISH 1914-04-22
40 Sicyos pachycarpus [Hawaiian Islands; U.S.A.; Hawaii Preserved_Specimen Rock, J.F.C. BISH
41 Sicyos pachycarpus Below Haleakala Ranch pipeline Preserved_Specimen Long trailing vine. Munro, G.C. 801 Maui BISH 1928-02-20
42 Sicyos pachycarpus Honolulu, Damon Tract coral rock, dry Preserved_Specimen Pearsall, G.A. 331 Oahu BISH 1959-04-03
43 Sicyos pachycarpus Kahului, along Koahumanu Avenue Preserved_Specimen Hobdy, R.W. 1367 Maui BISH 1982-04-22
44 Sicyos pachycarpus Koloa Distr., S side of road ca. 1 mile west of Poipu Beach Park Growing on wire fence. Preserved_Specimen Lamoureux, C.H. 2346 Kauai BISH 1963-04-13
45 Sicyos pachycarpus Mahana Preserved_Specimen Munro, G.C. 108 Lanai BISH 1913-10-20
46 Sicyos pachycarpus East Maui, Hana District, Nuu Preserved_Specimen Vine climbing over 8 m Prosopis trees; lvs membranous, dark green above, pale green below. Male fl white, female fl white, utricles green then black. Stem 5-angled, herbaceous, green. St.John, H. 26771 Maui BISH 1971-03-28
47 Sicyos pachycarpus Waimea District. Just outside the town of Hanapepe, along road following the Hanapepe River Large, scandent vine growing over largely introduced vegetation Preserved_Specimen Gustafson, R. 3577 Kauai BISH 1989-03-06
48 Sicyos pachycarpus Preserved_Specimen very lobed leaves Kepler, A.K. 9218 Laysan Island BISH 1992-04-14
49 Sicyos pachycarpus Kaena Preserved_Specimen Munro, G.C. 338 Lanai BISH 1914-04-22
50 Sicyos pachycarpus Kanaio, S slope of Haleakala Preserved_Specimen Forbes, C.N. 1621 Maui BISH 1920-03-03
51 Sicyos pachycarpus West side of coral plains at Ewa, near Barber's Point, ocean side of Malakole Rd. near Camp Malakole Military Reservation Preserved_Specimen Nagata, K.M. 1656 Oahu BISH 1978-06-21
52 Sicyos pachycarpus Laysan Preserved_Specimen White blossom Bryan, W.A. Laysan Island BISH 1911-05-01
53 Sicyos pachycarpus In two gullies S of Puu Kaua Preserved_Specimen Vines carpeting forest. Kikukawa, H. Oahu BISH 1985-05-08
54 Sicyos pachycarpus Laysan Preserved_Specimen white blossom Bryan, W.A. Laysan Island BISH 1911-05-01
55 Sicyos pachycarpus East Molokai, Kamiloloa Preserved_Specimen Hobdy, R.W. 2355 Molokai BISH 1985-04-17
56 Sicyos pachycarpus Near Sisal Preserved_Specimen Forbes, C.N. 1751 Oahu BISH 1912-02-16
57 Sicyos pachycarpus Basin, seaward between Middle and East Palm Valleys Preserved_Specimen Yen, D.E. 1006 Nihoa BISH 1969-05-01
58 Sicyos pachycarpus Waianae Mts., Palikea trail Preserved_Specimen Stone, B.C. 3475 Oahu BISH 1960-05-05
59 Sicyos pachycarpus Kamiloloa Preserved_Specimen Misaki, E.T. 7 Molokai BISH 1985-01-19
60 Sicyos pachycarpus Ewa, coral flats Preserved_Specimen Woolford, H.A. 410 Oahu BISH 1957-01-05
61 Sicyos pachycarpus Mahana Preserved_Specimen Munro, G.C. 171 Lanai BISH 1913-10-20
62 Sicyos pachycarpus Kaena Preserved_Specimen Munro, G.C. 90 Oahu BISH 1914-04-22
63 Sicyos pachycarpus Mountains, E end Preserved_Specimen Forbes, C.N. 278 Lanai BISH 1913-06-01
64 Sicyos pachycarpus Pohakea Pass Firebreak Trail; Ekahanui Gulch; Waianae Range Preserved_Specimen Kikudome, G.Y. 437 Oahu BISH 1951-05-27
65 Sicyos pachycarpus Palehua. Waianae Range Preserved_Specimen Kerr, M. 21443 Oahu BISH 1947-04-13
66 Sicyos pachycarpus Kalulu Preserved_Specimen Munro, G.C. 535 Lanai BISH 1916-03-24
67 Sicyos pachycarpus Kealia trail Preserved_Specimen Takeuchi, W.N. 61 Oahu BISH 1984-02-10
68 Sicyos pachycarpus Ewa Flats, Pohakupuna Rd Preserved_Specimen Woolford, H.A. 275 Oahu BISH 1957-01-13
69 Sicyos pachycarpus Basin, seaward between Middle and East Palm Valleys Preserved_Specimen Yen, D.E. 1006 Nihoa BISH 1969-05-01
70 Sicyos pachycarpus Kealia trail Preserved_Specimen Welton, P. 711 Oahu BISH 1991-05-11
71 Sicyos pachycarpus Kanepuu Preserved_Specimen Munro, G.C. 10 Lanai BISH 1927-05-28
72 Sicyos pachycarpus Coral plain below Ewa & Sisal Preserved_Specimen Forbes, C.N. 2323 Oahu BISH 1916-03-14
73 Sicyos pachycarpus Waianae Preserved_Specimen Takeuchi, W.N. 69 Oahu BISH 1984-02-20
74 Sicyos pachycarpus Coral plain below Ewa & Sisal Preserved_Specimen Forbes, C.N. 2323 Oahu BISH 1916-03-14
75 Sicyos pachycarpus Kanepuu Preserved_Specimen Munro, G.C. 10 Lanai BISH 1927-05-28
76 Sicyos pachycarpus Kaena Preserved_Specimen Munro, G.C. 90 Lanai BISH 1914-04-22
77 Sicyos pachycarpus Nanakuli Valley below Mauna Kapu Preserved_Specimen Degener, O. 17415 Oahu BISH 1935-12-01
78 Sicyos pachycarpus Coral plain below Ewa & Sisal Preserved_Specimen Forbes, C.N. 2323 Oahu BISH 1916-03-14
79 Sicyos pachycarpus East Maui, Waialoa, along Copp road Preserved_Specimen Hobdy, R.W. 4369 Maui BISH 2014-06-23
80 Sicyos pachycarpus Barbers Pt Preserved_Specimen Pearsall, G.A. 275 Oahu BISH 1950-06-02
81 Sicyos pachycarpus Kamokuiki Valley, between Puuiki and Puu Kamoahanui Preserved_Specimen Degener, O. 18035 Oahu BISH 1933-04-12
82 Sicyos pachycarpus Kamiloloa Preserved_Specimen Misaki, E.T. 6 Molokai BISH 1985-03-19
83 Sicyos pachycarpus Laysan Preserved_Specimen Herbst, D.R. 9063 Laysan Island BISH 1988-06-25
84 Sicyos pachycarpus West Maui, Maunahooma Preserved_Specimen Forbes, C.N. 11 Maui BISH 1910-05-01
85 Sicyos pachycarpus Preserved_Specimen Kepler, A.K. 9214 Laysan Island BISH 1992-04-14
86 Sicyos pachycarpus Waianae Mts., Keeau Valley Vining over flowers and fruit Preserved_Specimen Perlman, S. 5214 Oahu BISH 1987-02-10
87 Sicyos pachycarpus Kanaio, S slope of Haleakala Preserved_Specimen Forbes, C.N. 1621 Maui BISH 1920-03-03
88 Sicyos pachycarpus 3rd small valley NE of Palikea Preserved_Specimen Degener, O. 18036 Oahu BISH 1932-09-18
89 Sicyos pachycarpus Kamao Preserved_Specimen Munro, G.C. 418 Lanai BISH 1929-05-16
90 Sicyos pachycarpus Kalaupapa Beach Preserved_Specimen Rock, J.F.C. 14061 Molokai BISH 1918-05-24
91 Sicyos pachycarpus Coral plain below Ewa & Sisal Preserved_Specimen Forbes, C.N. 2323 Oahu BISH 1916-03-14
92 Sicyos pachycarpus Palikea Gulch, near Puu Pane, off firebreak road Preserved_Specimen Obata, J.K. 86633 Oahu BISH 1986-04-20
93 Sicyos pachycarpus Waiopai, S slope of Haleakala Preserved_Specimen Forbes, C.N. 1851 Maui BISH 1920-03-05
94 Sicyos pachycarpus Makua Valley; 1/8 mile east of Makua; Waianae Mountains Preserved_Specimen Carr, G.D. 918 Oahu BISH 1976-05-09
95 Sicyos pachycarpus [Hawaiian Islands; U.S.A.; Hawaii Preserved_Specimen Rock, J.F.C. BISH
96 Sicyos pachycarpus [Hawaiian Islands; U.S.A.; Hawaii Preserved_Specimen Rock, J.F.C. BISH
97 Sicyos pachycarpus Kamao Preserved_Specimen Munro, G.C. 418 Lanai BISH 1929-05-16
98 Sicyos pachycarpus Pohakea Pass, Ekahami Gulch, Waianae Range, firebreak trail Preserved_Specimen Kikudome, G.Y. 438 Oahu BISH 1951-05-27
99 Sicyos pachycarpus Preserved_Specimen Kepler, A.K. 9215 Laysan Island BISH 1992-04-14
100 Sicyos pachycarpus Preserved_Specimen Kepler, A.K. 9222 Laysan Island BISH 1992-04-16
101 Sicyos pachycarpus Preserved_Specimen Kepler, A.K. 9211 Laysan Island BISH 1992-04-14
102 Sicyos pachycarpus Preserved_Specimen Kepler, A.K. 9213 Laysan Island BISH 1992-04-14
103 Sicyos pachycarpus Preserved_Specimen Kepler, A.K. 9212 Laysan Island BISH 1992-04-14
104 Sicyos pachycarpus Huehue Preserved_Specimen Meebold, A. Hawaii BISH 1932-05-01
105 Sicyos pachycarpus Makena qd, Auwahi Preserved_Specimen Medeiros, A.C. Maui BISH 1984-01-01
106 Sicyos pachycarpus 26 degrees N, 172 degrees W Preserved_Specimen Kepler, A.K. Laysan Island BISH 1992-04-16
107 Sicyos pachycarpus Laysan Preserved_Specimen Bryan, W.A. Laysan Island BISH 1903-04-01
108 Sicyos pachycarpus Palehua. Waianae Range Preserved_Specimen Kerr, M. Oahu BISH 1947-04-13
109 Sicyos pachycarpus Ewa Flats, Pohakupuna Rd Preserved_Specimen Woolford, H.A. Oahu BISH 1957-01-13
110 Sicyos pachycarpus Laysan Preserved_Specimen Bryan, W.A. Laysan Island BISH 1903-04-01
111 Sicyos pachycarpus Kealia Salt Marsh Preserved_Specimen Funk, E.J. Maui BISH 1990-02-15
112 Sicyos pachycarpus Waialua Preserved_Specimen Caum, E.L. Oahu BISH 1946-03-12
113 Sicyos pachycarpus Maui Preserved_Specimen Clay, H.F. Maui BISH 1954-05-16
114 Sicyos pachycarpus Laysan Preserved_Specimen Beardsley, J.W. Laysan Island BISH 1962-06-15
115 Sicyos pachycarpus Barbers Point Preserved_Specimen Caum, E.L. Oahu BISH 1946-03-12
116 Sicyos pachycarpus Nihoa Preserved_Specimen Beardsley, J.W. Nihoa BISH 1962-06-10
117 Sicyos pachycarpus Maui Preserved_Specimen Clay, H.F. Maui BISH 1954-05-16
118 Sicyos pachycarpus Ewa, coral flats Preserved_Specimen Woolford, H.A. Oahu BISH 1957-03-11
119 Sicyos pachycarpus Waialua Preserved_Specimen Oahu BISH 1954-03-29
120 Sicyos pachycarpus [Hawaiian Islands; U.S.A.; Hawaii Preserved_Specimen Rock, J.F.C. 2661 BISH
121 Sicyos pachycarpus [Hawaiian Islands; U.S.A.; Hawaii Preserved_Specimen Rock, J.F.C. 2675 BISH
122 Sicyos pachycarpus East Maui, Makawao District, Kanaio Preserved_Specimen Holt, R.A. 152 Maui BISH 1982-03-23
123 Sicyos pachycarpus Waianae Mts., Honoliuli FR, Huliwai to Ekahanui gulches, ridge Preserved_Specimen Kikudome, G.Y. 275 Oahu BISH 1951-05-27
124 Sicyos pachycarpus Waiopai, S slope of Haleakala Preserved_Specimen Forbes, C.N. 1851 Maui BISH 1920-03-05
125 Sicyos pachycarpus [Hawaiian Islands; U.S.A.; Hawaii Preserved_Specimen Rock, J.F.C. BISH
126 Sicyos pachycarpus Kalaupapa Beach Preserved_Specimen Rock, J.F.C. 14061 Molokai BISH 1918-05-24
127 Sicyos pachycarpus unknown Preserved_Specimen Obata, J.K. BISH
128 Sicyos pachycarpus Waianae Mts., Haleiwa-Ekahanui gulch Preserved_Specimen Kikudome, G.Y. 275 Oahu BISH 1951-05-27
129 Sicyos pachycarpus Kamao Preserved_Specimen Munro, G.C. 418 Lanai BISH 1929-05-16
130 Sicyos pachycarpus Kalulu Preserved_Specimen Munro, G.C. 535 Lanai BISH 1916-03-24
131 Sicyos pachycarpus Kalaupapa Beach Preserved_Specimen Rock, J.F.C. 14061 Molokai BISH 1918-05-24
132 Sicyos pachycarpus Kanepuu Preserved_Specimen Munro, G.C. 10 Lanai BISH 1927-05-28
133 Sicyos pachycarpus NW of summit Young plants coming up through Lantana patch under a Kiawe tree. Preserved_Specimen Hobdy, R.W. 1250 Kahoolawe BISH 1982-02-18
134 Sicyos pachycarpus East Maui, Crater road near mile marker 1. Disturbed residential roadside. Preserved_Specimen 4 sheets, male and female flowers Welton, P. 1715 Maui BISH 1993-04-01
135 Sicyos pachycarpus near Makua Cave, Waianae climbing over Leucaena leucocephala Preserved_Specimen climbing; fruit pubescent Nagata, K.M. 3208 Oahu BISH 1985-03-17
136 Sicyos pachycarpus Waianae Kai Forest Reserve, Kamileunu Ridge near Puu Kepauala common Preserved_Specimen vine with white flowers Herbst, D.R. 5765 Oahu BISH 1976-03-14
137 Sicyos pachycarpus Lualualei steep rocky banks in Sapindus-Antidesma forest Preserved_Specimen Takeuchi, W.N. 3313 Oahu BISH 1987-01-31
138 Sicyos pachycarpus East Maui, Makawao District, Along the road behind La Perouse Bay In kiawe behind beach. Preserved_Specimen Vines. Corn, C. Maui BISH 1979-05-12
139 Sicyos pachycarpus Md. gulch S of Pohakea Pass, Waimea Mts, Halona, Lualualei For. Res., Lualualei Climbing on bushes, open slope. Preserved_Specimen St.John, H. 13136 Oahu BISH 1933-05-12
140 Sicyos pachycarpus Waianae Mts.; Honouliuli. Firebreak trail Growing on Grevillea robusta. Preserved_Specimen White flowers; vine 12 m. long. Chock, A.K. 238 Oahu BISH 1951-05-28
141 Sicyos pachycarpus Main ridge between Huliwai and Ekahanui gulches; Waianae Mountains; Honouliuli Large colony; climbing on Grevillea; Psidium; and Lantana. Preserved_Specimen Kikudome, G.Y. 435 Oahu BISH 1951-05-27
142 Sicyos pachycarpus Koloa District. Hanapepe, Papalu gulch, north facing slopes east of K. R. Wood cliff side property. Dodonaea viscosa mixed lowland dry shrubland w/ Heteropogon contortus, Sicyos pachycarpus, Doryopteris decora, Peperomia leptostachya, Plumbago zeylanica, Panicum konaense. Preserved_Specimen Vine, scandent along understory and climbig into canopy, common, in flower and fruit, Wood, K.R. 19307 Kauai BISH 2023-05-13
143 Sicyos pachycarpus Waianae Mts.; Honouliuli. Firebreak trail Growing on Grevillea robusta. Preserved_Specimen White flowers; vine 12 m. long. Chock, A.K. 238 Oahu BISH 1951-05-28
144 Sicyos pachycarpus East Maui, Hana District, Auwahi dry forest on aa Preserved_Specimen Vine with yellow-wh. fls., on aa Webster, G.L. 13887 Maui BISH 1968-06-25
145 Sicyos pachycarpus Makaha valley, E side on Leucaena Preserved_Specimen climbing; flowers white; fruit green, brown when ripe Davis, J. 103 Oahu BISH 1979-05-12
146 Sicyos pachycarpus W side under Scaevola Preserved_Specimen vine. Tendrils 2-3 branched, stems smooth, leaves papillosa Daniel, C.W. 8 Laysan Island BISH 1959-07-24
147 Sicyos pachycarpus Northwestern Hawaiian Islands East side of the lagoon ca. 200 m S of tide guage. Assoc. with Eragrostis & Ipomoea. Preserved_Specimen Vine; flrs w/white petals; frt green w/dark green protuberences. Conant, S. 166 Laysan Island BISH 1983-07-24
148 Sicyos pachycarpus NE slope of Mt. Kaala, 2nd gulch E of Kaupakuhale on bushes Preserved_Specimen vine, climbing St.John, H. 13183 Oahu BISH 1933-05-14
149 Sicyos pachycarpus Kalamaula In kiawe-Lantana forest. Preserved_Specimen Hobdy, R.W. 2379 Molokai BISH 1985-04-24
150 Sicyos pachycarpus East Maui, Hana District, Auwahi dry forest on aa Preserved_Specimen Vine with yellow-wh. fls. Webster, G.L. 13887 Maui BISH 1968-06-25
151 Sicyos pachycarpus Waianae Mountains; Honouliuli Forest Reserve; Pohakea Pass Trailing on bushes. Growing with Sicyos pachycarpus Hook. & Arnott. Preserved_Specimen Vine 20+ ft. long; roots slender. St.John, H. 13164 Oahu BISH 1933-05-12
152 Sicyos pachycarpus Mokuleia, Waialua, between Camp Erdman and Quarry over Leucaena from the talus to the highway Preserved_Specimen climbing Nagata, K.M. 2056 Oahu BISH 1980-04-12
153 Sicyos pachycarpus Kau On lava Preserved_Specimen Rock, J.F.C. Hawaii BISH 1911-04-01
154 Sicyos pachycarpus Kalamaula In kiawe-Lantana forest. Preserved_Specimen Hobdy, R.W. 2377 Molokai BISH 1985-04-24
155 Sicyos pachycarpus Waianae Mts., below ridge trail to Palikea, Waianae side, 1/4 mi. from radar station dry slope Preserved_Specimen vine, flowers white Cowan, R.S. 918 Oahu BISH 1948-03-27
156 Sicyos pachycarpus Ewa District. Pualii gulch, near Pohakea pass Degraded Acacia koa- Metrosideros mesic forest. With Aleurites moluccana, Xylosma hawaiiense, Myrsine lessertiana, Urera glabra, Antidesma platyphylla, Charpentiera obovata, Planchonella sandwicensis, Pisonia, Pipturus albidus, Tectaria gaudichaudi Preserved_Specimen Vine with flowers and fruit Perlman, S. 23555 Oahu BISH 2013-06-20
157 Sicyos pachycarpus Apua On slope of pali near kukui tree. Preserved_Specimen Fls green-white, 5 petals; climbing to 8 ft high. Fagerlund, G.O. 544 Hawaii BISH 1943-05-10
158 Sicyos pachycarpus East Maui, Nuu along road along road on rock wall. Disturbed roadside Preserved_Specimen male and female flower Welton, P. 1706 Maui BISH 1993-02-18
159 Sicyos pachycarpus East Maui, Makawao District, Kanaio Climbing over Leucaena leucocephala. Preserved_Specimen Vine. Fruits glabrous, numerous in globose heads 2 cm diam. Beak 1-2 mm. Stems pilose when young, glabrous w/ age. Leaves coarsely serrate, 5-angled to 5-lobed, pilose-papilose beneath. Male infl. sparsely glandular-pilose, 9-13 cm long, the racemes densely glandular-pilose. Male flrs w/ perianth lobes 3 mm, staminal column 1 mm. Female flrs w/ perianth lobes 1-1.5 mm. Nagata, K.M. 3191 Maui BISH 1985-03-11
160 Sicyos pachycarpus above Puu Olai, Makena Kiawe forests Preserved_Specimen Hobdy, R.W. 1370 Maui BISH 1982-04-25
161 Sicyos pachycarpus Kalamaula In kiawe-Lantana forest. Preserved_Specimen Hobdy, R.W. 2380 Molokai BISH 1985-04-24
162 Sicyos pachycarpus SE shore of lagoon over Cyperus with Eragrostis, Scaevola, and Ipomoea Preserved_Specimen vine with yellowish-white flowers Young, A.L. 156 Laysan Island BISH 1964-09-17
163 Sicyos pachycarpus Ekahanui Gulch, Honouliuli, Waianae Mts on dead tree trunk Preserved_Specimen Kikudome, G.Y. 383 Oahu BISH 1951-02-01
164 Sicyos pachycarpus along trail just N of Mauna Kapu, Waianae Mts with Bidens torta, Schinus, Psidium guajava, Acacia koa on ridge crest Preserved_Specimen Nagata, K.M. 3445 Oahu BISH 1986-02-22
165 Sicyos pachycarpus East Maui, Wailea, S of Polo Beach in Kiawe above road Preserved_Specimen Hobdy, R.W. 1371 Maui BISH 1982-04-25
166 Sicyos pachycarpus Waianae Mts.; Honouliuli. Firebreak trail Growing on Grevillea robusta. Preserved_Specimen White flowers; vine 12 m. long. Chock, A.K. 238 Oahu BISH 1951-05-28
167 Sicyos pachycarpus near Kawailoa Camp/Dump over haole koa Preserved_Specimen Takeuchi, W.N. 2133 Oahu BISH 1985-04-08
168 Sicyos pachycarpus Waianae, Keawaula, E of Farrington Hwy Locally abundant east of Farrington Hwy. Preserved_Specimen Leaves variable entire to deeple lobed. Nagata, K.M. 2043 Oahu BISH 1980-03-17
169 Sicyos pachycarpus N Kona, 28 mi S of Waimea, on Hwy 190 aa lava; with Nototrichium sandwicense, Diospyros ferrea sandwicensis, Chenopodium oahuense, Planchonella, Argemone; off roadside embankment Preserved_Specimen trailing vine Gustafson, R. 1233 Hawaii BISH 1979-06-19
170 Sicyos pachycarpus N Kona, Kaupulehu Preserve at mile 28 along Hawaii Belt Road 10 mi E of Kailua-Kona dry forest over aa lava with Diospyros sandwicensis, Colubrina oppositifolia, Santalum paniculatum, Nototrichium; scrambling over rocks and fountain grass Preserved_Specimen vine, leaves mottled green. Male and female flowers green and white. Fruit green, drying black when ripe. Lorence, D.H. 7991 Hawaii BISH 1997-05-15
171 Sicyos pachycarpus North Kona, Kaupulehu Dry weed lava slope 3 miles east of Huehue Preserved_Specimen Vine climbing 3 m. high; flowers yellow green Webster, G.L. 1837 Hawaii BISH 1948-06-23
172 Sicyos pachycarpus Ewa, Campbell Industrial Park, next to cement quarry In Prosopis forest. Preserved_Specimen Climbing over fallen Prosopis log. Flowering profusely. Leaves more deeply dissected nearer the apices Char, W.P. 81.005 Oahu BISH 1981-01-26
173 Sicyos pachycarpus East Maui, Crater road near mile marker 1. Disturbed residential roadside Preserved_Specimen male and female flowers Welton, P. 1716 Maui BISH 1993-04-20
174 Sicyos pachycarpus Mokuleia, bottom of Kealia Trail canopy Preserved_Specimen densely twining Conant, P. Oahu BISH 1992-11-20
175 Sicyos pachycarpus Slopes below Auahi Forest, S slope of Haleakala Common in open brushy land with a few scattered trees and large shrubs on aa lava. Preserved_Specimen Vine with leaves varying from angular to digitate on same plant. Fosberg, F.R. 48342 Maui BISH 1967-03-25
176 Sicyos pachycarpus East Maui at Nuu Bay in Kaupo Disturbed coastal lowland. Preserved_Specimen male and female flowers, fruit Welton, P. 1712 Maui BISH 1993-03-12
177 Sicyos pachycarpus Former Naval Air Station Barbers Point, Northern Trap & Skeet Range In Prosopis forest with Leucaena leucocephala, Panicum maximum, and Asystasia gangetica. Preserved_Specimen Entire leaf form. Vines growing up a Prosopis pallida 2.5 m above ground BISH staff 31 Oahu BISH 2021-04-30
178 Sicyos pachycarpus E of Farrington Hwy., Keawaula, Waianae on Leucaena leucocephala Preserved_Specimen Nagata, K.M. 2035 Oahu BISH 1980-02-17
179 Sicyos pachycarpus Honouliuli contour trail below Puu Kaua sunny grassy forest Preserved_Specimen Degener, O. 12751 Oahu BISH 1940-03-20
180 Sicyos pachycarpus Waianae Mts., Mauna Kapu, below summit edge of talus slope from road Preserved_Specimen Caum, E.L. Oahu BISH 1944-04-24
181 Sicyos pachycarpus SW of summit In a Kiawe - Lantana thicket. Preserved_Specimen Hobdy, R.W. 1307 Kahoolawe BISH 1982-03-12
182 Sicyos pachycarpus N end of lagoon over Pluchea shrubs Preserved_Specimen coarse vine with yellowish green flowers Herbst, D.R. 6519 Laysan Island BISH 1980-07-13
183 Sicyos pachycarpus Koloa Distr., S side of road ca. 1 mile west of Poipu Beach Park Growing on wire fence. Preserved_Specimen Lamoureux, C.H. 2346 Kauai BISH 1963-04-13
184 Sicyos pachycarpus SW of summit In a Kiawe - Lantana thicket. Preserved_Specimen Hobdy, R.W. 1305 Kahoolawe BISH 1982-03-12
185 Sicyos pachycarpus East Maui, Crater road near mile marker 1. Disturbed residential roadside. Preserved_Specimen 4 sheets, male and female flowers Welton, P. 1716 Maui BISH 1993-04-20
186 Sicyos pachycarpus below Mamahaloa Hwy., 2.4 mi from Huehue Ranch gate, Waimea, N Kona on Lantana, old aa flow; with Diospyros, Canthium, Pennisetum Preserved_Specimen climbing Nagata, K.M. 1752 Hawaii BISH 1978-07-28
187 Sicyos pachycarpus Waimea District, Queen's Pond area of Barking Sands Growing over Leucaena shrubs. Preserved_Specimen Vine. Herbst, D.R. 5691 Kauai BISH 1976-02-26
188 Sicyos pachycarpus SE shore of lagoon over Cyperus with Eragrostis, Scaevola, and Ipomoea Preserved_Specimen vine with yellowish-white flowers Young, A.L. 156 Laysan Island BISH 1964-09-17
189 Sicyos pachycarpus Former Naval Air Station Barbers Point, Northern Trap & Skeet Range, Outplanting site Growing on dead Prosops pallida in a burned area Southwest of San Juacinto St and Lamalle. With Solanum seaforthianum (779230), Leucaena leucocephala, Asystasia gangetica, Cenchrus ciliaris. Preserved_Specimen BISH staff 12 Oahu BISH 2021-04-29
190 Sicyos pachycarpus Makena In Kiawe forest above road. Preserved_Specimen Hobdy, R.W. 1369 Maui BISH 1982-04-25
191 Sicyos pachycarpus Waialua District, land of Kaena, Manini Pali area Common. Preserved_Specimen Annual vine. Herbst, D.R. 5844 Oahu BISH 1976-04-25
192 Sicyos pachycarpus East Maui, along Pililani highway in Kanaio. Dry lowland shrubland. Preserved_Specimen 3 sheets, male and female flowers, fruit Welton, P. 1909 Maui BISH 1996-03-24
193 Sicyos pachycarpus Kealia trail, bluffs behind Dillingham Airfield open clearing near top, over Panicum maximun Preserved_Specimen Takeuchi, W.N. 59 Oahu BISH 1984-02-10
194 Sicyos pachycarpus Waianae Mts.; Honouliuli. Firebreak trail Growing on Psidium guajava; in Grevillea robusta forest. Preserved_Specimen Kikudome, G.Y. 237 Oahu BISH 1951-05-28
195 Sicyos pachycarpus Kamakou Preserve, Upper Kupai'a Gulch, East Molokai Montane mesic or dry Metrosideros forest. Preserved_Specimen Vine climbing on shrubs. Anderson, S.J. Molokai BISH 1985-09-25
196 Sicyos pachycarpus Molokai Project - Cross-transect 3, large gulch nw of jeep trail, east Kamiloloa In shade, gulch bottom. Wiliwili-kiawe forest, koa-haole subcanopy layer. Preserved_Specimen Seedlings only. (Leaf shape similar to Herbst 2944 diposited at HAW) Char, W.P. 82.033 Molokai BISH 1982-11-27
197 Sicyos pachycarpus E of Farrington Hwy., Keawaula, Waianae on Leucaena Preserved_Specimen climbing Nagata, K.M. 2042 Oahu BISH 1980-03-06
198 Sicyos pachycarpus along Farrington Hwy., near Makaha Bridge 3-B, Keaau, Waianae drainage basin, sunny; on Prosopis, Leucaena Preserved_Specimen climbing on Prosopis and Leucaena. Nagata, K.M. 2041 Oahu BISH 1980-03-06
199 Sicyos pachycarpus Waianae Mts.; Honouliuli. Firebreak trail Growing on dead Acacia koa log. Preserved_Specimen Vine 8 m. long with green mature fruits. Chock, A.K. 240 Oahu BISH 1951-05-28
200 Sicyos pachycarpus Manini Gulch, Kaena, Waialua over Leucaena near base of cliffs Preserved_Specimen vine Nagata, K.M. 1627 Oahu BISH 1978-04-25
201 Sicyos pachycarpus Kaupulehu FR, Mamalahoa Hwy. over shrubs and trees Preserved_Specimen vine Herbst, D.R. 5296 Hawaii BISH 1975-05-02
202 Sicyos pachycarpus Waianae Mts, mouth of Kealia Gulch (behind Dillingham airfield) on south-facing slope Summer deciduous dry forest on cliffs. Associated with Erythrina, Reynoldsia, Rauwolfia, Diospyros, Dodonaea, Schiedea. Preserved_Specimen Vine. Flower white. Gagné, W.C. 671 Oahu BISH 1973-04-21
203 Sicyos pachycarpus NW of summit Young plants coming up through Lantana patch under a Kiawe tree. Preserved_Specimen Hobdy, R.W. 1251 Kahoolawe BISH 1982-02-18
204 Sicyos pachycarpus S side, ~50 m from lagoon sandy Preserved_Specimen prostrate Tsuda, R.T. 502 Laysan Island BISH 1963-12-05
205 Sicyos pachycarpus East Maui, Makawao District, Kanaio Natural Area Reserve Growing near the waterline road. Preserved_Specimen Oppenheimer, H.L. 49903 Maui BISH 1999-04-10
206 Sicyos pachycarpus Former Naval Air Station Barbers Point, Northern Trap & Skeet Range, Outplanting site Recently burned area. Growing on large Prosopis palida with ground cover of Cenchrus ciliaris, Megathyrsus maximus. Other seedlings of Sicyos around. Area shaded by canopy. Preserved_Specimen Vine growing 2 m above ground. Flowers white. BISH staff 13 Oahu BISH 2021-04-29
207 Sicyos pachycarpus East Maui, Kula district, c. 2 km from Keokea along road towards Kahului Gentle slope, W aspect. Clay loam on basalt Acacia mearnsii thicket edge. Frequent. Preserved_Specimen Climber. Stems to 5 m. Telford, I.R.H. 10233 Maui BISH 1986-05-10
208 Sicyos pachycarpus Northwestern Hawaiian Islands In shady moist mud/sand soil in a Coconut grove. Preserved_Specimen Vine; flrs yellow;frt green to brown (ripe). Conant, S. 159 Laysan Island BISH 1983-07-22
209 Sicyos pachycarpus Waianae Range, Kealia Trail, lower part of trail From near sea level in disturbed areas with Leucaena and Prosopis to Sapindus-dominated forest remains at ca. 100-150 m s.m. (collecting site for voucher). Preserved_Specimen Climbing vine with pistillate and staminate flowers, overgrowing even Leucaena and Prosopis. Kiehn, M. 9010312 Oahu BISH 2009-01-03
210 Sicyos pachycarpus NE slope of Mt. Kaala, 2nd gulch E of Puu Kaupakuhale on bushes Preserved_Specimen climbing vine St.John, H. 13183 Oahu BISH 1933-05-14
211 Sicyos pachycarpus Koloa Distr., S side of road ca. 1 mile west of Poipu Beach Park Growing on wire fence. Preserved_Specimen Lamoureux, C.H. 2346 Kauai BISH 1963-04-13
212 Sicyos pachycarpus W Maui, N of Honolowai disturbed area at edge of pineapple field Preserved_Specimen annual vine; flowers white, unisexual Annable, C.R. 3918 Maui BISH 1999-02-16
213 Sicyos pachycarpus Middle Valley (upper) In Ilima. Preserved_Specimen Vine Dried plants. Yen, D.E. 1006 Nihoa BISH 1969-05-01
214 Sicyos pachycarpus SW of summit In a Kiawe - Lantana thicket. Preserved_Specimen Hobdy, R.W. 1309 Kahoolawe BISH 1982-03-12
215 Sicyos pachycarpus near Kawailoa Camp/Dump over haole koa Preserved_Specimen Takeuchi, W.N. 2133 Oahu BISH 1985-04-08
216 Sicyos pachycarpus Makolelau Gulch Remnant of mesic native forest. Preserved_Specimen Vine. Flower white. Gagné, W.C. 676 Molokai BISH 1973-05-15
217 Sicyos pachycarpus Devil's slide scree, with deep soil, with Chenopodium oahuense Preserved_Specimen vine Conant, S. 113 Nihoa BISH 1983-04-30
218 Sicyos pachycarpus Gulch SW of Dillingham Ranch, Mokuleia In shade. Preserved_Specimen Degener, O. 20616 Oahu BISH 1950-04-23
219 Sicyos pachycarpus Mahaulepu, Kawailoa Bay, ~40 yds. behind beach Leucaena Preserved_Specimen Flynn, T.W. 802 Kauai BISH 1984-03-14
220 Sicyos pachycarpus Waianae Range, Honouliuli, Puu Kanehoa dry brushy slope Preserved_Specimen clambering vine with yellow flowers Webster, G.L. 1647 Oahu BISH 1948-06-13
221 Sicyos pachycarpus along Farrington Hwy., near Makaha Bridge 3-B, Keaau, Waianae sunny, drainage basin of intermittent stream; on Prosopis pallida, Leucaena leucocephala Preserved_Specimen climbing on Prosopis and Leucaena. Nagata, K.M. 2041 Oahu BISH 1980-03-06
222 Sicyos pachycarpus East Maui, Hana District, S slope of Haleakala, Auwahi Forest On rough, aa lava beds with scattered trees and shrubs. Very common, forming large patches climbing over scrub. Preserved_Specimen Extensive vine, flowers white fruit green. Fosberg, F.R. 55443 Maui BISH 1974-04-14
223 Sicyos pachycarpus SW of summit In a Kiawe - Lantana thicket. Preserved_Specimen Hobdy, R.W. 1308 Kahoolawe BISH 1982-03-12
224 Sicyos pachycarpus S side, ~50 m from lagoon sandy Preserved_Specimen prostrate Tsuda, R.T. 502 Laysan Island BISH 1963-12-05
225 Sicyos pachycarpus Makakupaia (Govt. Sect) Along road up to forest reserve, in a small gully. Found in Dodonaea scrub forming dense mats. Preserved_Specimen Hobdy, R.W. 2305 Molokai BISH 1985-03-20
226 Sicyos pachycarpus E of Farrington Hwy., Keaau, Waianae climbing on dense thickets of Leucaena leucocephala Preserved_Specimen leaves entire on seedlings, becoming shallowly to deeply 5-lobed on mature plants to 13 x 16 cm on petioles of 9 cm x 4 mm Nagata, K.M. 2040 Oahu BISH 1980-02-17
227 Sicyos pachycarpus Waianae Mts, mouth of Kealia Gulch (behind Dillingham airfield) on south-facing slope Summer deciduous dry forest on cliffs. Associated with Erythrina, Reynoldsia, Rauwolfia, Diospyros, Dodonaea, Schiedea. Preserved_Specimen Vine. Flower white. Gagné, W.C. 671 Oahu BISH 1973-04-21
228 Sicyos pachycarpus Former Naval Air Station Barbers Point, Northern Trap & Skeet Range, Outplanting site With Panicum maximum, Physalis angulata (779237), Asystasia gangetica, and Verbesina encelioides. Under Prosopis pallida. Preserved_Specimen BISH staff 19 Oahu BISH 2021-04-29
229 Sicyos pachycarpus Kealia trail, behind Dillingham Airfield over Leucaena Preserved_Specimen liane Takeuchi, W.N. 58 Oahu BISH 1984-02-10
230 Sicyos pachycarpus below Ewa and Sisal coral plain Preserved_Specimen Forbes, C.N. 2323 Oahu BISH 1916-03-14
231 Sicyos pachycarpus Puu Piimoe, Kanaio, National Guard target range, E side of cinder cone Dodonaea viscosa, Chenopodium oahuense, Myoporum sandwicense, Rauvolfia sandwicensis, Rhynchelytrum repens, Lipochaeta lavarum, Bidens pilosa, Nesoluma polynesicum, Nothocestrum latifolium, Ricinus communis, Doryopteris decipiens Preserved_Specimen Wood, K.R. 5138 Maui BISH 1996-04-24
232 Sicyos pachycarpus Palikea Gulch near Puu Pane, firebreak road, Waianae Mts on road cut, over shrubs Preserved_Specimen Obata, J.K. 77314 Oahu BISH 1977-04-17
233 Sicyos pachycarpus Land of Makakupaia. Burned area of pines. Just inside forest reserve. Preserved_Specimen Huge masses draping plants, corollas white, leaf lobing polymorphic as is fruit pubescence. Wagner, W.L. 6674 Molokai BISH 1992-01-18
234 Sicyos pachycarpus Polihale State Park, mouth of Kaulaula valley, south of Haeleele. Secondary non-native vegetation dominated by Leucaena leucocephala and Prosopis pallida. w/ some relic trees of Erythrina sandwicensis, Psydrax odorata Preserved_Specimen Vine, blanketing volcanic basalt boulders, in flower and immature fruit Wood, K.R. 15851 Kauai BISH 2014-02-27
235 Sicyos pachycarpus near Makua Cave, Waianae climbing over Leucaena leucocephala Preserved_Specimen climbing; fruit pubescent Nagata, K.M. 3208 Oahu BISH 1985-03-17
236 Sicyos pachycarpus Roadside in Kanaio area. S slope of Haleakala Leucaena and Lantana. Found occasionally as far east as Nuu. Preserved_Specimen Vine Nagata, K.M. 1944 Maui BISH 1979-06-07
237 Sicyos pachycarpus Waianae Mts.; Honouliuli. Firebreak trail Growing on Psidium guajava with Grevillea robusta. Preserved_Specimen White flowers. vine 5 m. long; with several auxilliary branches Chock, A.K. 235 Oahu BISH 1951-05-28
238 Sicyos pachycarpus Waimea, Puu Ki-Waialae Trail along Mokihana Ridge with Metrosideros, Styphelia, Dodonaea, Acacia, Wilkesia, Psidium, Grevillea Preserved_Specimen vine; stems pale green; leaves medium to dark dull green above, much paler green below; male flowers; corolla pale yellow-green, five-lobed Flynn, T.W. 4438 Kauai BISH 1991-02-07
239 Sicyos pachycarpus above Puu Olai, Makena kiawe forests Preserved_Specimen Hobdy, R.W. 1368 Maui BISH 1982-04-25
240 Sicyos pachycarpus Koloa, Lawai Valley, Allerton Gardens, along Stream by 'cut flower' garden over shrubs Preserved_Specimen vine; flowers sweetly fragrant (10 am). Monecious, flowers with greenish white corollas Lorence, D.H. 7361 Kauai BISH 1993-02-18
241 Sicyos pachycarpus Kealia trail, bluffs mauka of Dillingham Air Field over Leucaena Preserved_Specimen A liana Takeuchi, W.N. 58 Oahu BISH 1984-02-10
242 Sicyos pachycarpus Honouliuli, N fork of valley, E of Palikea, Waianae Mts open woods Preserved_Specimen vine St.John, H. 10388 Oahu BISH 1930-02-23
243 Sicyos pachycarpus East Maui, Ulupalakua Gentle slope, W aspect. Clay loam on basalt Acacia mearnsii thicket edge. Frequent. Preserved_Specimen Climber. Stems to 5 m. Telford, I.R.H. 10230 Maui BISH 1986-05-10
244 Sicyos pachycarpus Kalamaula In kiawe-Lantana forest. Preserved_Specimen Hobdy, R.W. 2378 Molokai BISH 1985-04-24
245 Sicyos pachycarpus Waimea District; Koaie Canyon between 1 and 2 miles above Lonomea Camp Herbaceous climber in Lantana thicket. Preserved_Specimen Leaves dark dull green above; dull below; stems whitish-green; male flowers white at base with dark green apex. Flynn, T.W. 453 Kauai BISH 1991-04-16
246 Sicyos pachycarpus Former Naval Air Station Barbers Point, Northern Trap & Skeet Range, Outplanting site Growing on dead Prosops pallida in a burned area Southwest of San Juacinto St and Lamalle. With Solanum seaforthianum (779230), Leucaena leucocephala, Asystasia gangetica, Cenchrus ciliaris. Preserved_Specimen BISH staff 12 Oahu BISH 2021-04-29
247 Sicyos pachycarpus Waianae arid roadside Preserved_Specimen Degener, O. 18391 Oahu BISH 1932-04-10
248 Sicyos pachycarpus Kauooiokunui Gulch open kukui forest Preserved_Specimen Degener, O. 12752 Oahu BISH 1940-04-11
249 Sicyos pachycarpus Waianae Mts.; Honouliuli. Firebreak Trail. Located near Chock & Kikudome 235 Vine growing on Psidium guajava with roots in ground. Preserved_Specimen White flowers. Chock, A.K. 236 Oahu BISH 1951-05-28
250 Sicyos pachycarpus Turned east onto jeep road just south of Manawainui Gulch. Passed Homelani Cemetary, on jeep road to Molokai Forest Reserve. On south side of road. Kalamaula area Vine in Prosopis grove. Dry. Preserved_Specimen Herbst, D.R. 2944 Molokai BISH 1973-04-03
251 Sicyos pachycarpus Waimea District, Queen's Pond area of Barking Sands Growing over Leucaena shrubs. Preserved_Specimen Vine. Herbst, D.R. 5691 Kauai BISH 1976-02-26
252 Sicyos pachycarpus Waianae Mts.; Honouliuli. Firebreak trail Growing on Grevillea robusta. Preserved_Specimen White flowers; vine 12 m. long. Chock, A.K. 238 Oahu BISH 1951-05-28
253 Sicyos pachycarpus W. slope of Tanager Peak Growing on Chenopodium sandwicense. Preserved_Specimen Vine Christophersen, E. 9 Nihoa BISH 1924-07-12
254 Sicyos pachycarpus Waianae Mts., Pohakea Pass in Eucalyptus, Grevillea Preserved_Specimen Herbst, D.R. 463 Oahu BISH 1967-04-29
255 Sicyos pachycarpus Lualualei, end of Dent St among Leucaena leucocephala Preserved_Specimen twining Lamoureux, C.H. 3842 Oahu BISH 1967-05-21
256 Sicyos pachycarpus Keaau, Waianae District. E of Farrington Hwy In dense thickets of Leucaena leucocephala. Preserved_Specimen Leaves variable entire to deeple lobed. Nagata, K.M. 2043 Oahu BISH 1980-03-17
257 Sicyos pachycarpus SE shore of lagoon over Cyperus with Eragrostis, Scaevola, and Ipomoea Preserved_Specimen vine with yellowish-white flowers Young, A.L. 156 Laysan Island BISH 1964-09-17
258 Sicyos pachycarpus Auwahi, E. Maui sparse dryland forest; locally common on ravine slopes Preserved_Specimen Female clusters 1 cm diam. Flowers white. Male inflorescences on peduncles 5 cm long Nagata, K.M. 3876 Oahu BISH 1988-08-16
259 Sicyos pachycarpus Kalamaula, NW of Puu Luahine On fences or climbing trees, in pasture with Prosopis, Leucaena, and relict Erythrina Preserved_Specimen Occasional annual vines Oppenheimer, H.L. 40901 Molokai BISH 2009-04-01
260 Sicyos pachycarpus Makaha valley, E side on Leucaena Preserved_Specimen climbing; flowers white; fruit green, brown when ripe Davis, J. 103 Oahu BISH 1979-05-12
261 Sicyos pachycarpus S end dark sand Preserved_Specimen prostrate vine, 10 cm; tendrils 2-3 branched, stems smooth, leaves papillose Daniel, C.W. 18 Laysan Island BISH 1959-07-25
262 Sicyos pachycarpus Kaupo, E of Pohakuloa growing on shrubs. Preserved_Specimen Low vines Hobdy, R.W. 1031 Maui BISH 1981-03-18
263 Sicyos pachycarpus Near summit ( lower block 5 of Forestry Tree Planting Project) Dense thicket of Lantana and Kiawe Preserved_Specimen Trailing vine Hobdy, R.W. 1965 Kahoolawe BISH 1984-02-16
264 Sicyos pachycarpus ridge S of Huliwai Gulch, Waianae Mts., Honouliu on trees Preserved_Specimen Kikudome, G.Y. 434 Oahu BISH 1951-05-27
265 Sicyos pachycarpus E of Farrington Hwy., Keaau, Waianae climbing on dense thickets of Leucaena leucocephala Preserved_Specimen leaves entire on seedlings, becoming shallowly to deeply 5-lobed on mature plants to 13 x 16 cm on petioles of 9 cm x 4 mm Nagata, K.M. 2040 Oahu BISH 1980-02-17
266 Sicyos pachycarpus Kaluakauila Valley, Keawaula, Waianae Mts on Grevillea trunks, partly sunny, dry, well sloped Preserved_Specimen Obata, J.K. 284 Oahu BISH 1975-04-06
267 Sicyos pachycarpus Ekahanui Gulch; 50 yds. from ridge divide; S of ridge between this and Huliwai Gulch; Waianae Mountains; Honouliuli Over Lantana brush. Preserved_Specimen Kikudome, G.Y. 382 Oahu BISH 1951-02-01
268 Sicyos pachycarpus East Maui, along Piilani highway in Kanaio dry lowland shrubland Preserved_Specimen male and female flowers, fruit Welton, P. 1909 Maui BISH 1996-03-24
269 Sicyos pachycarpus Lualualei steep rocky banks in Sapindus, Antidesma forest Preserved_Specimen Takeuchi, W.N. 3313 Oahu BISH 1987-01-31
270 Sicyos pachycarpus W. slope of Tanager Peak Growing on Chenopodium sandwicense. Preserved_Specimen Christophersen, E. 9 Nihoa BISH 1924-07-12
271 Sicyos pachycarpus Ahihi-Kinau Natural Area Reserve, Ahihi Bay at the N end of the reserve Pockets of vegetation near the seashore amongst a'a lava about 200 years old.Vine with quadrangular stems; growing on stumps of Prosopis. Clusters of female flowers; panicles of male. Fruit globose aggregation of beaked fruits. Preserved_Specimen Abbott, I.A. 91.8 Maui BISH 1991-03-05
272 Sicyos pachycarpus Makakupaia (Govt. Sect) Along road up to forest reserve, in a small gully. Found in Dodonaea scrub forming dense mats. Preserved_Specimen Hobdy, R.W. 2305 Molokai BISH 1985-03-20
273 Sicyos pachycarpus W Maui, Manawainui Plant Sanctuary, Lower Hanaula Twining over shrubs on sunny gulch bottom. Preserved_Specimen Hobdy, R.W. 1060 Maui BISH 1981-04-16
274 Sicyos pachycarpus North Kona; Kapulehu Dry weed lava slope 3 miles east of Huehue. Preserved_Specimen Vine climbing 3 m. high; flowers yellow green. Webster, G.L. 1837 Hawaii BISH 1948-06-23
275 Sicyos pachycarpus above Nuu rough lava flow Preserved_Specimen Forbes, C.N. 1906 Maui BISH 1920-03-09
276 Sicyos pachycarpus Between Barber's Point and Pearl Harbor arid, fossil reef back of beach Preserved_Specimen leaves variable Degener, O. 18394 Oahu BISH 1932-05-08
277 Sicyos pachycarpus NW part in bunchgrass Preserved_Specimen vine Herbst, D.R. 6533 Laysan Island BISH 1980-07-14
278 Sicyos pachycarpus Waianae Mts.; Honouliuli. Firebreak trail Growing on Psidium guajava; in Grevillea robusta forest. Preserved_Specimen Vine 2 m. long. Chock, A.K. 237 Oahu BISH 1951-05-28
279 Sicyos pachycarpus behind Kaena Pt on Leucaena shrubs; talus slopes Preserved_Specimen Nagata, K.M. 3819 Oahu BISH 1988-02-27
280 Sicyos pachycarpus Onini Gulch Vine in gulch bottom Preserved_Specimen Vine in gulch bottom. Bishop, L.E. 1690 Molokai BISH 1973-01-21
281 Sicyos pachycarpus near Kawailoa Camp/Dump draped over haole koa Preserved_Specimen Takeuchi, W.N. 2133 Oahu BISH 1985-04-08
282 Sicyos pachycarpus Naval Reservation, Lualualei Valley, end of Dent Street Climbing on koa haole . Arid, w/introduced plants: Bidens pilosa, Cenchrus echinatus, Schinus terebinthifolius. Preserved_Specimen Flowers, buds, mature fruit present. Herbst, D.R. 492 Oahu BISH 1967-05-21
283 Sicyos pachycarpus Waianae Mts.; Honouliuli. Firebreak trail Growing on Grevillea robusta. Preserved_Specimen Vine with green mature fruits. Chock, A.K. 239 Oahu BISH 1951-05-28
284 Sicyos pachycarpus W Maui, Lahaina District, Papalaua headwaters, W fork Climbing adjacent vegetation in open areas of Mesic Forest & Shrubland. Vegetation: Metrosideros, Diospyros, Myrsine, Pleomele, Alyxia, Dodonaea, Wikstroemia, Leptecophylla, Dryopteris, Doryopteris, Doodia, Microlepia, Asplenium, Smilax, Bidens, Eragrostis, Carex, Cenchrus, Panicum, Luzula, Erigeron, Adiantum, Ageratina, Lantana. Preserved_Specimen Local annual vines to 4m long. Differs from the description in the sparsely pubescent stems (vs. glabrous), typical in many Maui populations. Oppenheimer, H.L. 40803 Maui BISH 2008-04-09
285 Sicyos pachycarpus Former Naval Air Station Barbers Point, Northern Trap & Skeet Range In Prosopis forest with Leucaena leucocephala, Panicum maximum, and Asystasia gangetica. Preserved_Specimen Palmate leaf form. Vines growing up a Prosopis pallida 2.5 m above ground BISH staff 32 Oahu BISH 2021-04-30
286 Sicyos pachycarpus Kalamaula In kiawe-Lantana forest. Preserved_Specimen Hobdy, R.W. 2376 Molokai BISH 1985-04-24
287 Sicyos pachycarpus SW of summit In a Kiawe - Lantana thicket. Preserved_Specimen Hobdy, R.W. 1306 Kahoolawe BISH 1982-03-12
288 Sicyos pachycarpus 2nd gulch of Puu Kaupakuhale, NE slope of Puu Kaala, Waianae Mts., Mokuleia bushes Preserved_Specimen vine, climbing St.John, H. 13183 Oahu BISH 1933-05-14
289 Sicyos pachycarpus Upper Makakilo, Kalo‘i Gulch, in gulch bottom Disturbed shady gulch floor with overstory of Aleurites moluccana, Leucaena leucocephala, Schinus terebinthifolius, understory dominated by Panicum maximum, with Hyptis pectinata and Desmanthus pernambucanus. Groves of Erythina sandwicensis scattered in vicinity. Preserved_Specimen Native vine rampantly spreading at this time of year over the Guinea grass. Imada, C. 200430 Oahu BISH 2004-03-30
290 Sicyos pachycarpus Ewa Beach Marina, inland from Batis swamp dry branches, with wild tobacco and popolo berries Preserved_Specimen Funk, E.J. 603 Oahu BISH 1990-05-01
291 Sicyos pachycarpus Mahaulepu, Kawailoa Bay sand dune, with Casuarina equisetifolia, Prosopis Preserved_Specimen trailing, stems 2 m Telford, I.R.H. 10244 Kauai BISH 1986-06-02
292 Sicyos pachycarpus Kalamaula In kiawe-Lantana forest. Preserved_Specimen Hobdy, R.W. 2375 Molokai BISH 1985-04-24
293 Sicyos pachycarpus Near summit ( lower block 5 of Forestry Tree Planting Project) Dense thicket of Lantana and Kiawe Preserved_Specimen Trailing vine Hobdy, R.W. 1965 Kahoolawe BISH 1984-02-16
294 Sicyos pachycarpus NE of Pohaka Pass trees below cliff Preserved_Specimen climbing Degener, O. 21362 Oahu BISH 1949-07-17
295 Sicyos pachycarpus Mahaulepu, Kawailoa Bay, ~40 yds. behind beach Leucaena Preserved_Specimen Flynn, T.W. 802 Kauai BISH 1984-03-14
296 Sicyos pachycarpus E of Farrington Hwy., Keawaula, Waianae climbing on Leucaena. Preserved_Specimen climbing Nagata, K.M. 2042 Oahu BISH 1980-03-06
297 Sicyos pachycarpus main ridge S of Kanehoa and Huliwai Gulch, Honouliuli, Waianae Mts on trees Preserved_Specimen climbing Kikudome, G.Y. 433 Oahu BISH 1951-05-27
298 Sicyos pachycarpus Makolelau Gulch In Metrosideros - Dodonaea viscosa - Styphelia lowland dry shrubland with Alyxia; Styphelia; Melicope; Pleomele; Scaevola; Pteridium; Sida; Osteomeles; Schiedea lydgatei; Bobea; Diospyros; Coprosma; Wikstroemia; Neraudia sericea; weeds: Ageratina adenophora Preserved_Specimen Plants with Flowers and fruits. Perlman, S. 11986 Molokai BISH 1991-06-12
299 Sicyos pachycarpus Makaleha Valley, Mokuleia over brush and weeds Preserved_Specimen vine, climbing St.John, H. 25961 Oahu BISH 1956-04-15
300 Sicyos pachycarpus Kamakou Preserve, Upper Kupai'a Gulch, East Molokai. Montane mesic or dry Metrosideros forest. Preserved_Specimen Vine climbing on shrubs. Anderson, S.J. Molokai BISH 1985-09-25
301 Sicyos pachycarpus Along the crater road in upper Kula Growing in a thicket of Acacia decurrens Preserved_Specimen Hobdy, R.W. 1380 Maui BISH 1982-05-03
302 Sicyos pachycarpus Lahaina District, Kauaula valley, S slope, below Helu summit, W Maui Diospyros sandwicensis-Metrosideros mesic forest. With Nestegis sandwicensis, Pisonia, Zanthoxylum hawaiiense, Cyrtandra grayi, Corprosma foliosa, Cheirodendron trigynum, Dodonaea viscosa, Polyscias oahuensis, Myrsine lanaiensis, Sophora chrysophylla Preserved_Specimen Vine with flowers and immature fruit. Perlman, S. 22840 Maui BISH 2012-04-18
303 Sicyos pachycarpus Koloa, Eleele, top of bluff, SE of Hanapepe River, behind School with Leucaena; on fence Preserved_Specimen vine; flowers pale green, fruit green. Lorence, D.H. 5830 Kauai BISH 1988-02-27
304 Sicyos pachycarpus Koloa Distr., S side of road ca. 1 mile west of Poipu Beach Park Growing on wire fence. Preserved_Specimen Lamoureux, C.H. 2346 Kauai BISH 1963-04-13
305 Sicyos pachycarpus East Maui, Makawao District, Along the road behind La Perouse Bay In kiawe behind beach. Preserved_Specimen Vines. Corn, C. Maui BISH 1979-05-12
306 Sicyos pachycarpus Makaha. Makaha Valley; Waianae Mts Climbing in Psidium guajava thicket. Preserved_Specimen St.John, H. 25121 Oahu BISH 1953-05-03
307 Sicyos pachycarpus Vacant lot between Malakole Ave. & deep draft harbor. Campbell Industrial Park, Ewa Common vine climbing on Leucaena. Wasteland of Leucaena, Cenchrus ciliaris & Panicum maximum. Preserved_Specimen Nagata, K.M. 3950 Oahu BISH 1989-01-28
308 Sicyos pachycarpus East Maui, Mauka of Puu o kali hill Preserved_Specimen Fruits very dark when mature. Hobdy, R.W. 2063 Maui BISH 1984-04-19
309 Sicyos pachycarpus Ewa Plantation cultivated land Preserved_Specimen Menardi, J.B. Oahu BISH 1952-01-23
310 Sicyos pachycarpus East Maui, Hana District, Nuu Preserved_Specimen Vine climbing over 8 m Prosopis trees; lvs membranous, dark green above, pale green below. Male fl white, female fl white, utricles green then black. Stem 5-angled, herbaceous, green. St.John, H. 26771 Maui BISH 1971-03-28
311 Sicyos pachycarpus gulch below Palikea, Honouliuli, Waianae Range Preserved_Specimen Lus. thin, pale, soft; flowers white. Fruit dark green Cowan, R.S. 628 Oahu BISH 1947-06-15
312 Sicyos pachycarpus Ewa, coral flats coral flats Preserved_Specimen Woolford, H.A. Oahu BISH 1957-03-11
313 Sicyos pachycarpus E of Farrington Hwy., Keawaula, Waianae on Leucaena leucocephala Preserved_Specimen Nagata, K.M. 2035 Oahu BISH 1980-02-17
314 Sicyos pachycarpus East Maui, Lava flows above Kihei, below Puu o Kali cone Abundant in lava flows from 500 to 1400 ft. Preserved_Specimen Hobdy, R.W. 1004 Maui BISH 1981-03-05
315 Sicyos pachycarpus Mountains near Koele Preserved_Specimen Large vine. Forbes, C.N. 95 Lanai BISH 1913-06-01
316 Sicyos pachycarpus S end Preserved_Specimen prostrate vine; flowers white; tendrils 3-branched, stems smooth Daniel, C.W. 33 Laysan Island BISH 1959-07-26
317 Sicyos pachycarpus Mountains near Koele Preserved_Specimen Large vine. Forbes, C.N. 95 Lanai BISH 1913-06-01
318 Sicyos pachycarpus Laysan Preserved_Specimen Herbst, D.R. 9057 Laysan Island BISH 1988-06-19
319 Sicyos pachycarpus Mokuleia, slopes of Kaala Preserved_Specimen Forbes, C.N. 1783 Oahu BISH 1912-04-26
320 Sicyos pachycarpus Ewa Beach and Quarry Preserved_Specimen Christophersen, W. 141 Oahu BISH 1982-01-30
321 Sicyos pachycarpus East Maui, Makawao District, along road behind La Perouse Bay In kiawe behind the beach Preserved_Specimen Corn, C. Maui BISH 1979-05-12
322 Sicyos pachycarpus Kealia trail Preserved_Specimen Takeuchi, W.N. 60 Oahu BISH 1984-02-10
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